Anne Shirley, an orphan, is fostered by farmer Matthew Cuthbert and his sister Marilla, who were expecting a boy to be sent them to help with their farm work. They accept Anne, who quickly endears herself to them and to the local villagers.

Honoka Inoueas Anne Shirley (voice)
Aya Nakamuraas Marilla Cuthbert (voice)
Naoya Miyaseas Gilbert Blythe (voice)
Yume Miyamotoas Diana Barry (voice)
Kimiko Saitoas Rachel Lynde (voice)
Chie Nakamuraas Mrs. Spencer (voice)
Sayaka Senbongias Flora Jane Spencer (voice)
Seiko Tamuraas Mrs. Blewett (voice)
Akihiko Ishizumias Stationmaster (voice)
Rena Maedaas Eliza Barry (voice)
Michiru Oshima